Meadow Browns and Ringlets

I used to think that the Meadow Brown was probably the most common butterfly in East Lothian. I had memories of seeing hundreds of them on grasslands on the coast, along field edges and up in the hills. However, over the last few years I haven't seen many at all. Possibly I have been out and about at the wrong time, or maybe my memory is letting me down.

Last year numbers of Meadow Browns were lower than average, despite what seemed like favourable weather. However, as I commented previously the dry weather and easterly winds may have resulted in these lower numbers.

Looking at the figures that I have been gathering for the last 14 years, numbers seem to vary quite a bit from year to year and if anything have been showing a slight increase.

The number of the other grass-feeder, Ringlets also vary quite a bit from year to year, but they do seem to show a downward trend.

At home in the Borders I have certainly seen an increase in numbers of both species, but since we took over the place ten years ago, we have created a couple of areas of wild flower meadows and felled an area of Spruce woodland, replacing it with native broadleaf trees, with grass growing under them. I think I can put the increase in Meadow Browns and Ringlets here down to there being a lot more suitable habitat for them.

I had a look at my notes from when I had been surveying Northern Brown Argus in the valley above our house, but there was nothing conclusive as the numbers of Meadow Browns and Ringlet were really variable and they were based on only a couple of visits a year.

However, last year colleagues in East Lothian and the Butterfly Recorder in the Borders also commented that they had noticed a reduction in the number of Meadow Browns and Ringlets. This caused me to look for any other records I had.

I was able to look at the transect figures from Aberlady Local Nature Reserve going back to 2008. Both Meadow Brown and Ringlet numbers have been really variable, but both show quite an obvious down turn in numbers.

The other long-standing transect in East Lothian is at John Muir Country Park. It was set up in 1995, but unfortunately there are quite a number of years when it hasn't been walked. However, it shows a similar decline in numbers.

It is really interesting to note that Ringlets weren't recorded there until 1998. It is easy to forget that they only made their way up to East Lothian in the early 1980s.

I will certainly keep an eye on how numbers do both here in the Borders and in East Lothian over the next few years. I would be interested to hear how these two species are doing elsewhere across the UK.

Scottish Borders Butterflies 2021

Due to Covid travel restrictions our planned holiday to Spain was cancelled for the second year in a row. I always look forward to a trip abroad and the opportunity to see different butterflies.  On previous trips to Spain I have managed to see about 40 species of butterflies, so for the second year in a row my tally was going to be particularly low.

I am lucky that we have some land where we have done much to encourage butterflies and just a few hundred metres above our house there is an amazing valley where there are a lot of interesting species. There are a few interesting species that occur a few miles away, so I decided this year to see how many species I could find in the Scottish Borders.

The year started off well with a couple of Small Tortoiseshells, Aglais urticae,  visiting the garden in March and in April Peacocks, Aglais io, and Commas, Polygonia c-album,  joined them. Despite the good start to the season the weather was cold and wet later in the spring and I didn't find any caterpillars of these species, which is unusual. However, they obviously did manage to breed as there were more adults later in the summer, but not in as good numbers as normal.

I am always pleased to see the first Orange Tip, Anthocharis cardamines, of the season, as to me they mark the start of the butterfly season. This year they did well here and were seen in good numbers.

Of the other three species of whites, the Green-veined White, Pieris napi, didn't do as well as normal for some reason. Unusually, it was outnumbered by the Small White, Pieris rapae. Large Whites, Pieris brassicae, are never really numerous, but they were a regular visitor.

In early  June I drove over to Burnmouth to look for Small Blues, Cupido mimimus. Although there was a cool breeze coming in off the sea I still saw quite a few, along with Small Heaths, Coenonympha pamphilus, Small Coppers, Lycaena phlaeas, and Wall Browns, Lasiommata megera.

Back home and a walk up the valley added Northern Brown Argus, Aricia artaxerxes, and Small Pearl-bordered Fritillary, Clossiana selene, to the count along with loads of Small Heaths, Ringlets, Aphantpopus hyperantus, and Meadow Browns, Maniola jurtina.

Red Admirals, Vanessa atalanta, arrived in June and in September their numbers picked up as the new generation appeared. I was amazed how long they remained here with the last one I saw being on the 25th October, long after the first frosts.

At the beginning of July I headed towards Treepwood where I had seen old records of Large Heaths, Coenonympha tullia. It was difficult to find a suitable location amongst the fields grazed by sheep, but eventually I managed to find a boggy area with Cotton Grass growing. After much stumbling about I was delighted to find one Large Heath. I am not sure if numbers would have been higher earlier or later in the year, but hopefully they are doing well there.

In another visit up the valley above the house that month there were a few Dark Green Fritillaries, Argynnis aglaja, and loads of Small Skippers, Thymelicus sylvestris. Amongst them I spotted one Large Skipper, Ochlodes sylvanus.

Towards the end of July I went to Kelso to look for White-letter Hairstreaks, Satyrium w-album. Iain Cowe, the Borders butterfly recorder, had previously shown me how to find eggs on Elm trees, so I headed to the same spot. Within a few minutes I spotted some butterflies high in the trees and a little later one kindly flew down and landed on a branch next to me.

I was planning on a visit to a valley a few miles away to see Scotch Argus, Erebia aethiops, but the weather at the weekends was disappointingly poor. However, one lunchtime when I was walking around my meadow, I spotted a dark butterfly which turned out to be a beautiful fresh Scotch Argus. Of course, I didn't have my camera with me and I couldn't find it once I had run back to the house to get it! However, six days later I spotted a quite faded Scotch Argus in the meadow. Unfortunately, I will never know if it was the same butterfly!

So, I managed to see 23 species of butterflies in the Scottish Borders this year. I don't think there are many more species that I could have seen. Green Hairstreak, if I had been in the right place, possibly Grayling somewhere on the coast and Painted Lady, had there been more around this year. I was very pleased to have seen a Large Heath and White-letter Hairstreak - two butterflies I have never seen before.

East Lothian Butterflies 2017 - Part 2

Following on from my previous post here are the remaining species recorded in East Lothian in 2017.

Green Hairstreak, Callophrys rubi

The Green Hairstreak is probably very much under-recorded in East Lothian. There are old records from the Lammermuir Hills, but they have not been seen in those sites for many years. They have, however, been discovered in other areas of the Lammermuirs and in woodlands on the East Lothian side. They require blaeberries as food plants for the caterpillars, and everywhere we have found them is within a few feet of conifer trees. Numbers appeared to be down in 2016, but it is difficult to tell how they are doing without more specific monitoring.

Painted Lady, Vanessa cardui

Painted Lady numbers are very variable year on year, depending on many factors on their long migration from Africa to the UK. We don't ever see enormous numbers of them in East Lothian. The first record for East Lothian was on 5th May and it looks as though there was a better than average arrival of Painted Ladies. They didn't go on to produce a large second brood, presumably as a result of the summer weather.

Small Heath, Coenonympha pamphilus

The Small Heath confuses me a little! In the south of the UK there are two generations a year and up here we are supposed to get just one generation. However, I have noticed a big difference between sites in East Lothian. They are rarely seen beyond the end of June in John Muir Country Park, yet in other sites are seen through until the end of August. In 2017 the first record was on 6th May. They started off doing well, but their numbers were lower later in the season. We have been getting later records each year, but in 2017 we had a remarkably late record of one being seen on 20th October. I can only imagine this was from a second brood.

Common Blue, Polyommatus icarus

The Common Blue was seen in slightly lower numbers than normal in 2017, with sightings being lower than average towards the end of the season. The first Common Blue was recorded on 31st May and they had a fairly normal year with numbers just a bit down on the average.

Small Pearl Bordered Fritillary, Clossiana selene

The Small Pearl Bordered Fritillary is a very rare butterfly in East Lothian, only being recorded in a couple of places in the Lammermuir Hills and at John Muir Country Park. This year I only received one record on 7th June from John Muir Country Park.

Ringlet, Aphantpopus hyperantus

Ringlets did reasonably well in 2017. The first record received was on 14th June and their numbers built up quickly to a maximum, three weeks later and then back down to zero in the next five weeks.

Northern Brown Argus, Aricia artaxerxes

The Northern Brown Argus only occurs in a few small colonies in East Lothian. There is one very well established colony in the Lammermuirs, a very small and precarious colony on a golf course near Dunbar and possibly another couple of inaccessible sites where they occur. They tend not to travel far from their food plant, Rockrose, and therefore don't have much likelihood of spreading in East Lothian. The first record in 2017 was on 17th June and because we receive so few records it is difficult to assess how well they are doing.

Large Skipper, Thymelicus sylvestris

It was very pleasing to receive two records of Large Skippers in 2017. In 2014 we received our first record on the coast just inside East Lothian. We had expected them to continue along the coast, as many other species have, but despite many people searching there were no sightings in 2015 or 2016. Then on 17th June 2017 someone spotted them in two locations in the foothills of the Lammermuir Hills. They had caught us all off guard and sneaked west! I am sure there will be plenty of us out checking appropriate sites along the Lammermuirs in 2018!

Meadow Brown, Maniola jurtina

The Meadow Brown is normally the most numerous butterfly in East Lothian, but this year it was knocked back into four place. The first record we received was on 19th of June and this coincided with the start of the horrible rainy weather. It is interesting to see that numbers picked up to almost normal levels in August once the rain had stopped!

Small Skipper, Thymelicus sylvestris

The Small Skipper was first recorded in East Lothian in 2011 and since then it has been increasing in number and spreading across the county. This butterfly was first spotted at Aberlady bay, then in Saltoun Wood, other sites along the coast, Gifford, Linn Dean and Haddington to mention just a few. It would be interesting to know it manages to get to all these sites. Does it fly large distances, or is it that we just haven't noticed it in places between these sites?

Grayling, Hypparchia semele

The Grayling is only regularly seen in two or three sites in East Lothian. Unfortunately, one of those sites, Blindwells, is due to be developed and public access is being restricted. They are normally seen between mid-June to the end of August. The first record I received in 2017 was on 24th June and numbers seemed low this year.

Dark Green Fritillary, Argynnis aglaja

The first record in 2017 of a Dark Green Fritillary was on 1st July. Sadly, because their emergence was during the rainy weather 2017 was their worst year of the last five years. Numbers of ad hoc and transect records were both less than half what would normally be expected.

Sadly, 2017 was the first year that I haven't received a record for a Holly Blue since 2009. There was a colony around the Newhailes and Brunstane area, which sadly seems to have died out and each year I have received one or two records from around the Gullane/North Berwick area. I live in hope that there is a little colony hidden away somewhere in that area.

I am very grateful to all of those who have contributed butterfly records. It shouldn't be too long before we are seeing butterflies again in 2018!

Butterflies through time (1)

We only have 25 different species of butterflies here in East Lothian. However, thirty years ago there were only about half that number. I have always been interested in why we have recorded so many new species here in recent years. Is it because we are putting more effort into searching for the butterflies, or are these genuine increases in the range of these species?

Recently I was lucky enough to find a second-hand copy of "The Butterflies of Scotland" by George Thomson.  This was writen in 1980 and I was keen to see the distribution maps and compare them with those in the 1970 "Provisional Atlas of the Insects of the British Isles" and with the current distribution maps produced by Butterfly Conservation. However, I was delighted to discover that the author has also included the history of records for each species going back to the early 1800s.

I have been spending hours reading this book and learning about how the distribution of many species of butterflies has changed over the years. Some of what I have read confirms what people have told me but there were a few surprised in there too!


I will start with some now common butterflies. The Orange Tip, Anthocharis cardamines, is said to have been widespread throughout southern and east Scotland in the early 1800s and it remained so until the 1880s, but after that its range contracted to south west Scotland. In about 1950 it was considered to be advancing back towards the Lothians. The map in Butterflies of Scotland shows it occurring in South-west Scotland and the Highlands, with one or two records in between. My boss, who was a countryside ranger in the 1970s says he remembered them only being along the River Esk in the west of East Lothian. Since then they have spread and they are now commonly seen in the spring across much of East Lothian and Scotland.


The Peacock, Aglais io, was recorded in south west Scotland in the 1700s and it continued to be recorded there through the 1800s with sporadic records across Scotland. However, by the start of the 1900s it was becoming rarer. It is thought that small colonies persisted on the west coast of Scotland, but that other records from around Scotland were migrants. Records started to pick up again in the 1950s and since then it has continued to spread across Scotland. It wasn't until the 1980s that it was found in East Lothian.

Ringlets, Aphantopus hyperantus, were said to be abundant in East Lothian in 1852, but by the end of that century they were becoming scarce. The last record from East Lothian around that time was in 1928. It was recorded again in 1984 in the east of East Lothian and has since spread across the whole county, now being common and numerous.

There are records of the Comma, Polygonia c-album, occurring in Southern Scotland, and up to Fife in the 1850s, but after that its range contracted until in 1920s it was only regularly occurring in the southern half of England. From around the 1980s it underwent a rapid expansion and reached the Scottish Borders in the 1990s and was first recorded again in East Lothian in 2001. Since 2006 it has become established here and has slowly become more numerous.

So, three of these species were certainly in East Lothian in the early 1800s, and most likely the Peacock was also occasionally found here then, too. Now the question is not so much why they occur here now, but why they moved away and then returned. I will have a look at the other more recent arrivals in East Lothian and see if I can find information about the weather during this period. More to follow...

East Lothian Butterflies 2013

The weather in 2013 was a big improvement on the previous two years. Although the winter wasn’t particularly hard, it dragged on for a long time, and spring didn’t arrive until half way through April. This, and I think last year’s very poor weather, had an impact on the number of butterflies early in the year. Other than a couple of sightings in March, the first records of butterflies didn’t come in until 26th April!

However, from mid April until the end of August, the weather was reasonably warm, dry and sunny. It was never particularly hot, so there was a continuous source of food plants and the number of butterflies really picked up during July and August. The weather in September and October was rather disappointing, so we didn’t get the influx of butterflies migrating north, that we were hoping for. The first frosts arrived in early November, putting an end to the butterfly season.

Peacock, Aglais io

The first record I received this year was a Peacock, seen on North Berwick Law on the 1st March. There were a few seen throughout the spring, but the number seen dropped at the end of May. In August, though, there was a sudden explosion of Peacocks when the next generation appeared. Numbers through August were the highest I can remember seeing.

Peacock

Small Tortoiseshell, Aglais urticae

The first record this year of a Small Tortoiseshell in East Lothian was on 2nd March. After that they appeared regularly through to May. Like the Peacock their numbers picked up considerably with the new generation in July and August.

Small Tortoiseshell

Painted Lady, Vanessa cardui

One Painted Lady was recorded on 25th April, but it wasn’t until the end of June that more were seen having worked their way northwards through Europe. They weren’t seen up here in great numbers this year, though.

Painted Lady

Green-veined White, Pieris napi

The butterfly season kicked off properly with the first record of a Green-veined White on 26th April. Numbers were a little low, but they were seen regularly up until the middle of June. In July the second generation appeared with a vengeance. It seems that the weather conditions must have been perfect for them this year with numbers peaking around the middle of August.

Green-veined White

Small White, Pieris rapae

The first record of a Small White this year was on 29th April, which is more than a month later than last year and two months later than 2011. This just shows how their emergence is related to the weather. Like the Green-veined Whites, their numbers were low for the spring generation, but when the second generation started to appear at the beginning of August they were seen in far greater numbers than they have been for many years.

Small White

Comma, Polygonia c-album

The Comma is a butterfly that isn’t seen in great numbers here. It was first recorded in East Lothian in 2004 and is now seen all over the county, but rarely more than one at a time. The first one recorded here this year was on 30th April and a few records kept trickling in until the end of October. I find it odd that we never see it in greater numbers like the Small Tortoiseshell or Peacock with which it shares a similar life cycle and food plant.

Comma

Orange Tip, Anthocharis cardamines

The first Orange Tip seen this year was on 7th May. Their numbers were lower than average, but considering the rain and flooding we had last year this isn’t surprising. They don’t have a second summer generation like the other whites, but hopefully they should do well next year. Strangely, I didn’t see very many eggs or caterpillars, which are normally fairly easy to spot.

Orange Tip

Large White, Pieris brassicae

The first Large White was seen on 7th May. Generally we don’t see Large Whites in great numbers here, however this year, as with the other species from August onwards we had unusually high numbers of Large Whites. It wasn’t unusual to see more than 40 on a buddleia bush.

Large White

Speckled Wood, Pararge aegeria

Speckled Woods are butterflies that excite me a lot! In 2009 we received our first couple of records of them in East Lothian. The following year I found a small colony of them and since then they have increased in numbers year on year. The first record this year was on 14th May. After that records came in from all over the north and middle of the county. It seems that they first arrived here on the east coast having spread up from the Scottish Borders. This year they have worked their way almost along the entire length of the coast and up the River Tyne valley. It is amazing to see such a rapid spread of this species and next year I won’t be surprised if they are seen all over the county.

Speckled Wood

Holly Blue, Celastrina argiolus

I only received one record of a Holly Blue this year on the 21st May. This was sad after they had been seen in various locations over the previous two years. Hopefully this one sighting means that they are still clinging on at the long-established colony in the west of East Lothian.

Red Admiral, Vanessa atalanta

The first Red Admiral we saw this year was on May 22nd. Thereafter they were seen a few times, but compared with other butterflies this year, they weren't around in great numbers. I think the long winter was too much for any of them to survive and we didn't get very many making their way up from Europe.

Small Copper, Lyceana phlaeas

The weather this year seemed to suit Small Coppers with the first one being recorded on 30th May and thereafter higher than normal numbers being spotted. It was interesting to note that last year the majority of those seen were of the caeruleopunctata aberration, with a number of blue spots on their hind wings. However, this year very few aberrations were noted. It is interesting to speculate whether the aberration is a result of the climate, or quality of the caterpillar food plant, or for some other reason.

Small Copper

Wall Brown, Lamiommata megera

Like the Speckled Woods, Wall Browns were first recorded in East Lothian only a few years ago, but they don’t seem to have spread as quickly. We had a new record of them on North Berwick Law, so they have spread about half way along the coast in that time. The first record this year was on 30th May. They seem to occur mostly on the coast, but in August I saw one six kilometres inland at Woodhall Dean.

Wall Brown

Small Heath, Coenonympha pamphilus

The first Small Heath was recorded on 2nd June. It is a common butterfly along the coast in East Lothian and it is also found in the Lammermuir Hills. This year they appeared to do as well as ever.

Small Heath

Common Blue, Polyommatus icarus

The common Blue had a fairly normal year. I think that because they have come from eggs that were laid last year, they seem to be affected less by this year's good weather. The first Common Blue was seen on 16th June and they were on the wing until the middle of September.

Common Blue

Ringlet, Aphantopus hyperantus

Ringlets have a very short flight period. They occur in damp grassy areas and normally start to appear in late June, and this year they turned up right on cue on 21st June. The number of Ringlets on my transect this year was lower than normal, but that isn't surprising given that the area was flooded twice last year. At other sites around East Lothian there were good numbers of Ringlets.

Ringlet

Grayling, Hipparchia semele

The Grayling was restricted to a couple of very small sites in East Lothian, with the odd record of them being found at other coastal sites in the past, but these seem to be one-offs. The first Grayling recorded this year was on 25th June at one of the established sites. Last year a new colony was discovered on a re-landscaped mining spoil heap close to one of their other colonies. When I visited this colony in mid July I counted 80 individuals in a short visit, double the number that I saw last year.

Grayling

Meadow Brown, Maniola jurtina

Meadow Browns seemed to do very well this year. The first record was on 25th June and they were seen in good numbers up until the end of August. I only saw one on my transect this year, but again, this isn’t surprising given the flooding we had in that area last year.

Meadow Brown

Dark Green Fritillary, Argynnis aglaja

The Dark Green Fritillary is a lovely bright butterfly that occurs in many coastal areas and valleys in the Lammermuir Hills. The first one was seen in East Lothian this year on 30th June. They never occur in great numbers, and this year proved to be about average for them.

Dark Green Fritillary

Northern Brown Argus, Aricia artaxerxes

I am aware of four colonies of Northern Brown Argus in East Lothian, but I am sure there are probably more in the Lammermuir Hills. The colonies are very small, one being based around a patch of their food plant – the Rockrose, Helianthemum nummularium on a golf course and another on a small patch of Rockrose in a private garden.  I tend to go and specifically look for them where I know they occur and this year they seemed to be a little later than normal, but in reasonable numbers.

Northern Brown Argus

Small Pearl-bordered Fritillary, Boloria selene

Having visited a reserve in the Scottish Borders this year to see Small Pear-bordered Fritillaries, I was delighted to find them at a site just inside East Lothian a couple of weeks later. There have previously been a few unconfirmed sightings of Small Pearl-bordered Fritillaries in East Lothian, but this time I saw them long enough and took some pictures which confirmed their identification. It is funny how seeing a butterfly elsewhere can lead to identifying them in different sites. If I hadn’t seen those in the Scottish Borders I may have dismissed these as Dark Green Fritillaries and not looked any closer!

Small Pearl-bordered Fritillary

Small Skipper, Thymelicus sylvestris

Another very exciting record this year was the discovery of a number of Small Skippers at Aberlady Local Nature Reserve. There had been three isolated sightings of Small Skippers at two locations in East Lothian the previous two years, but on 12th July I received an excited phone call from a local volunteer who had found more than ten of them on a patch of thistles in the reserve.  She went on to find more at the reserve and I then received other records of them at three other locations within three kilometres of this site. It is great to hear that they are doing well and if we have good weather next year I am sure we will find them at various other locations.

Small Skipper

Clouded Yellow, Colias croceus

We only very rarely see Clouded Yellows here. I saw one thirteen years ago and a colleague saw one seven years ago. This year we received two records. The first was seen in North Berwick on 31st July and the following day we were told of one about 25 kilometres further along the coast in Musselburgh. I guess it must have been the same individual. In August and September high numbers arrived in southern England from the continent. They were seen laying eggs and I was hoping that we would be invaded by the next generation. Unfortunately the cold weather proved too much for them and they didn't venture up here!

All together this has been a fantastic year for butterflies. In total I received sightings of 23 different species, which I think must be a record for East Lothian. Only ten years ago eight of those species hadn't been recorded here. It is amazing that so many species continue to expand their range into East Lothian. We seem to be losing more and more habitat, and yet the butterflies seem to be better than ever. I wonder what the next new species will be. I can't wait for next year!